Gratiana spadicea (Klug, 1829)
Cassida spadicea Klug, 1829: 8; Boheman, 1854: 390, 1856: 128, 1862: 307; Burmeister, 1870: 280; Gemminger and Harold, 1876: 3658.
Odontionycha spadicea: Weise, 1896 c: 29.
Gratiana spadicea: Spaeth, 1913 b: 142, 1914 g: 121; Frers, 1925: 82 (imm. stages); Tenebaum, 1927: 37; Kvasina and Ponce de Leon, 1985: 9-15 (incl. fig., larva and bionomy), 1988: 1-6 (karyotype); Becker and Freiro-Costa, 1987: 195-205 (ootheca and biology); Albuquerque and Becker, 1986: 105-113 (biology); Buzzi, 1988: 574 (biology); Borowiec, 1996 a: 188, 1999 d: 375, 2002 a: 94, 2009 f: 672; Ponce de Leon et al., 1999: 1; Mediros and Moreira, 2002: 295-305 (biology), 2008: 210-224 (biology); Nogueira de Sa et al., 2004: 651 (bionomy).
Nuzonia spadicea: Blackwelder, 1946: 750.
Host plant: Solanaceae: Solanum sisymbriifolium (Frers, 1925; Albuquerque and Becker, 1986; Kvasina and Ponce de Leon, 1985, 1988, Ponce de Leon et al., 1999; Byrne et al., 2002, Medeiros and Moreira, 2002, 2008).
Distr.: Süd-Brasilien. N Argentina: Buenos Aires, Chaco de Santiago del Estero, Gran Chaco, Jujuy, Misiones, San Juan; Brazil: Mato Grosso, Paraná, Rio Grande do Sul, Sao Paulo; Paraguay: Asuncion, Col. Thomson, Paraguari; Uruguay: Montevideo.
figure: habitus in colour
previous species: pallidula